Sunday, November 03, 2019

Invasive species


An invasive species is a species that is not native to a specific location (an introduced species), and that has a tendency to spread to a degree believed to cause damage to the environment, human economy or human health. The term as most often used applies to introduced species that adversely affect the habitats and bioregions they invade economically, environmentally, or ecologically. 


Such species may be either plants or animals and may disrupt by dominating a region, wilderness areas, particular habitats, or wildland-urban interface land from loss or natural controls (such as predators or herbivores).  This includes plan species labeled as exotic pest plants and invasive exotics growing in native plant communities. The European Union defines "Invasive Alien Species" as those that are, firstly, outside their natural distribution area, and secondly, threaten biological diversity.


The term is also used by land managers, botanists, researchers, horticulturalists, conservationists, and the public for noxious weeds. The term "invasive" is often poorly defined or very subjective and some broaden the term to include indigenous or "native" species, that have colonized natural areas, for example, deer considered by some to be overpopulating their native zones and adjacent suburban gardens in the Northeastern and Pacific Coast regions of the United States.


The definition of "native" is also sometimes controversial. For example, the ancestors of Equus ferus (modern horses) evolved in North America and radiated to Eurasia before becoming locally extinct. Upon returning to North America in 1493 during their hominid-assited migration, it is debatable as to whether they were native or exotic to the continent of their evolutionary ancestors.


Notable examples of invasive plant species include the kudzu vine, Andean pampas grass, and yellow star-thistle. Animal examples include the New Zealand mud snail, feral pigs, European rabbits, grey squirrels, domestic cats, carp and ferrets. Invasion of long-established ecosystems by organisms from distant bio-regions is a natural phenomenon, but has been accelerated massively by humans, from their earliest migrations though to the age of discovery, and now international trade.


While all species compete to survive, invasive species appear to have specific traits or specific combinations of traits that allow them to outcompete native species. In some cases, the competition is about rates of growth and reproduction. In other cases, species interact with each other more directly. Researchers disagree about the usefulness of traits as invasiveness markers. One study found that of a list of invasive and noinvasive species, 86% of the invasive species could be identified from the traits alone.


Another study found invasive species tended to have only a small subset of the presumed traits and that many similar traits were found in noinvasive species, requiring other explanations. Common invasive species traits include the following: 

- Fast growth
- Rapid reproduction 
- High dispersal ability
- Phenotype plasticity
- Tolerance of a wide range of environmental conditions
- Ability to live off of a wide range of food types
- Association with humans
- Prior successful invasions


Typically, an introduced species must survive at low population densities before it becomes invasive in a new location. At low population densities, it can be difficult for the introduced species to reproduce and maintain itself in a new location, so a species might reach a location multiple times before it becomes established. Repeated patterns of human movement, such as ships sailing to and from ports or cars driving up a down highways offer repeated opportunities for establishment (also known as a high propagule pressure).


An introduced species might become invasive of it can outcompete native species for resources such as nutrients, light, physical space, water, or food. If these species evolved under great competition or predation, then the new environment may host fewer able competitors, allowing the invader to proliferate quickly. Ecosystems which are being used to their fullest capacity by native species can be modeled as zero-sum systems in which any gain for the invader is a loss for the native. However, such unilateral competitive superiority (and extinction of native species with increased populations of the invader) is not the rule. 


Invasive species often coexist with native species for an extended time, and gradually, the superior competitive ability of an invasive species becomes apparent as its population grows larger and denser and it adapts to its new location. An invasive species might be able to use resources that were previously unavailable to native species, such as deep water sources accessed by a long taproot, or an ability to live on previously uninhabited soil types. For example, barbed goatgrass was introduced to California on serpentine soils, which have low water-retention, low nutrient levels, a high magnesium/calcium ratio, and possible heavy metal toxicity. Plant populations on these soils tend to show low density, but goatgrass can form dense stands on these soils and crowd out native species that have adapted poorly to serpentine soils.


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